Dr. Benjamin Naumann


Universität Rostock

Institut für Biowissenschaften

Allgemeine & Spezielle Zoologie

Universitätsplatz 2, 18055 Rostock

Telefon: 0381 / 498-6267

Email: benjamin.naumann (at) uni-rostock.de

ResearchGate: Benjamin Naumann


The influence of developmental constraints on the evolution of direct development in frogs and other amphibians

Ecological developmental evolution (Eco-Evo-Devo) of aplocheiloid killifishes

Evolutionary and functional morphology of diplopod copulatory organs

The evolution and development (Evo-Devo) of cell differentiation and multicellularity in choanoflagellates and early-branching metazoans


2014 - 2018: PhD thesis at the Friedrich-Schiller-Universität Jena: “Head, trunk and in between – the evolution and development of muscles in the head/trunk interface of vertebrates”

2011 – 2014: Master of Science “Ecology, Evolution and Systematics” at the Friedrich-Schiller-Universität Jena, Thesis: “The development of the cucullaris and the hypobranchial muscles in the Mexican axolotl and the zebrafish”

2008 – 2011: Bachelor of Science “Biology” at the Friedrich-Schiller-Universität Jena, Thesis: “The development of the cucullaris muscle in the Mexican axolotl”


October 2021 – present: Scientific employee (Postdoc) at the Lehrstuhl für Allgemeine und Spezielle Zoologie, Universität Rostock

2018-2021: Scientific employee (Postdoc) at the Institut für Zoologie und Evolutionsforschung, Friedrich-Schiller-Universität, Jena

2014-2018: Scientific employee (PhD candidate) at the Institut für Zoologie und Evolutionsforschung, Friedrich-Schiller-Universität, Jena



Sachkova, M., Nordmann, E. L., Angel, J. J. S., Meeda, Y., Gorski, B., Naumann, B., ... & Burkhardt, P. (2021). Neuropeptide repertoire and 3D anatomy of the ctenophore nervous system. Current Biology.

Voigtlander, K., Burkhardt, U., Dannefeld, T., Heynen, E., Lindner, N., Moritz, L., Naumann, B., Orsakowsky, R., Sombke, A., Reip, H. (2021). The Myriapoda of the Schorfheide-Chorin UNESCO Biosphere Reserve in Brandenburg (Germany). Schubartiana 9, 13-37.


Naumann, B., Schweiger, S., Hammel, J. U., & Muller, H. (2020). Parallel evolution of direct development in frogs - Skin and thyroid gland development in African Squeaker Frogs (Anura: Arthroleptidae: Arthroleptis). Developmental Dynamics 250 (4), 584-600.

Levit, G., Hosfeld, U., Naumann, B., Lukas, P. & Olsson, L. (2020). The biogenetic law and the Gastraea theory: From Ernst Haeckel’s discoveries to contemporary views. The Journal of Experimental Zoology Part B 2021, 1-15.

Gohde, R.*, Naumann, B.*, Laundon, D., Imig, C., McDonald, K. L., Cooper, B. H., Varoqueaux, F., Fasshauer, D. & Burkhardt, P. (2020). Choanoflagellates and the ancestry of neurosecretory vesicles. Proceedings of the Royal Society B 376 (1821), 20190759. (*shared first authorship)

Schmidt, J. Schuff, M., Naumann, B. & Olsson, L. (2020). FoxN3 knockdown alters spatio-temporal expression patterns of genes involved in cranial cartilage and muscle development in the African clawed-frog, Xenopus laevis. Vertebrate Zoology, 70 (3), 455-472.

Naumann, B., Reip, H. S., Akkari, N., Neubert, D., Hammel, J. U. (2020). Inside the head of a cybertype – 3D reconstruction of the head muscles of Ommatoiulus avatar Akkari and Enghoff, 2015 (Diplopoda: Julidae) and insights on the head musculature of Juliformia. Zoological Journal of the Linnaean Society, 188 (4), 954-975.


Naumann, B., & Burkhardt, P. (2019). Spatial cell disparity in the colonial choanoflagellate Salpingoeca rosetta. Frontiers in Cell and Developmental Biology, 7, 231.

Naumann, B., Schmidt, J., & Olsson, L. (2019). FoxN3 is necessary for the development of the interatrial septum, the ventricular trabeculae and the muscles at the head/trunk interface in the African clawed frog, Xenopus laevis (Lissamphibia: Anura: Pipidae). Developmental Dynamics, 248 (5), 323-336.


Naumann, B., & Englert, C. (2018). Dispersion/reaggregation in early development of annual killifishes: Phylogenetic distribution and evolutionary significance of a unique feature. Developmental Biology, 442 (1), 69-79.

Naumann, B., & Olsson, L. (2018). Three‐dimensional reconstruction of the cranial and anterior spinal nerves in early tadpoles of Xenopus laevis (Pipidae, Anura). Journal of Comparative Neurology, 526 (5), 836-857.


Warth, P., Hilton, E. J., Naumann, B., Olsson, L., & Konstantinidis, P. (2017). Development of the muscles associated with the mandibular and hyoid arches in the Siberian sturgeon, Acipenser baerii (Acipenseriformes: Acipenseridae). Journal of Morphology,279 (2), 163-175.

Naumann, B., Warth, P., Olsson, L., & Konstantinidis, P. (2017). The development of the cucullaris muscle and the branchial musculature in the Longnose Gar (Lepisosteus osseus, Lepisosteiformes, Actinopterygii) and its implications for the evolution and development of the head/trunk interface in vertebrates. Evolution & Development, (19) 6, 263-276.

Schweiger, S., Naumann, B., Larson, J.G., Mockel, L. & Muller, H. (2017). Direct development in the African squeaker frogs (Anura: Arthroleptidae: Arthroleptis) reveals a mosaic of derived and plesiomorphic characters. Organisms, Diversity and Evolution, 17 (3), 693-707.


Warth, P., Konstantinidis, P., Hilton, E.J., Naumann, B. & Olsson, L. (2016). Development of the skull and pectoral girdle in Siberian sturgeon, Acipenser baerii, and Russian sturgeon, Acipenser gueldenstaedtii (Acipenseriformes: Acipenseridae). Journal of Morphology, 278 (3), 418-442.


Konstantinidis, P., Warth, P., Naumann, B., Metscher, B., Hilton, E.J., Olsson, L. (2015). The Developmental Pattern of the Musculature Associated with the Mandibular and Hyoid Arches in the Longnose Gar, Lepisosteus osseus (Actinopterygii, Ginglymodi, Lepiosteiformes). Copeia, 103 (4), 920-932.